100 The World of Biology (4)
    An introduction to biology.  Provides an overview of the study of life, including how biology affects our  daily lives, including such topics as biodiversity, genetic  engineering, and problems  associated with the expanding human population. Designed  for non-science majors. $50 lab fee. Offered every semester.
 117 Gardening and Plant  Propagation (2)
  An introduction to concepts  surrounding plant growth  and nutrition with a practical  application toward the maintenance of a  vegetable garden and experience working within the VWU greenhouse. Beyond the  garden, topics include basic plant tissue culture, grafting, cloning and  hydroponics. Offered each spring.
130 Principles of Biology I: Ecology and Evolution (4)
    An introduction to the biological sciences. Topics include ecology,  Darwinian evolution, population genetics, speciation, the origin of life, animal form and  function, animal behavior, and biological conservation. Designed for students  intending to pursue a major in biology, EES or ENVS. Offered every fall.
 132 Principles of Biology  II: Cell Biology  and Genetics (4)
    An introduction to the biological sciences, focused on topics  including biochemistry, cell structure &  metabolism, the cell cycle  & cancer biology, gene inheritance, gene expression, and explores  applications into biotechnology. Designed for students intending to pursue a  major in biology, biochemistry, or chemistry. $50 lab fee. Offered every  spring.
 133 Principles of Ecology  and Evolution (4)
    Introduction to the biological sciences with a focus on Darwinian evolution, population genetics, speciation, the origin of life,  animal form and function, animal behavior, ecology and biological conservation.  Designed for dual enrolled high school students only. Includes a laboratory  component. $50 lab fee. Offered intermittently.
150 Introduction to Marine Biology (4)
    An introduction to the organisms and communities of marine and estuarine areas.  Students examine the basic physical  and ecological processes that are pertinent to marine habitats and the  diversity of marine organisms and ecosystems. Includes field and laboratory  identification of local organisms and investigations of local field habitats.  Designed for non-science majors. $50 lab fee. Offered every fall.
 195 Pathogens, Parasites and Plagues  (4)
    Designed for none-majors, this course explores  the nature of human diseases  caused by bacteria,  viruses and other  parasitic organisms. Includes a review of entomology, the biology of parasites, and the essentials of disease transmission. This course focuses on  the techniques for eradication of disease in the U.S. and undeveloped nations.
 200 Principles of Biology  III: The Diversity  of Life (4)
    Completes the introduction to the  life sciences for Biology majors with a survey of all major lineages of  organisms on the planet, including bacteria  and archaea, fungi,  plants, protists, and animals. $50 lab fee. Prerequisite: BIO 130 or BIO 132 and  sophomore status or higher. Offered every fall.
201 Microbiology for the Health  Sciences (4)
    This course is intended  for those students  enrolled in the joint Nursing  Program with the Sentara College  of Health Sciences and is an introduction to the  science of the biology of microorganisms as applied to health care settings.  $50 lab fee.
    Prerequisite: CHEM 130. Offered  intermittently.
202 Human  Nutrition (4)
    This course is designed to  introduce processes by which the human body obtains nutrients from food to  maintain structural and functional integrity. We explore dietary  sources, intake levels and the requirement of major nutrients, role of nutrition  in growth and human health, and the role of diet in chronic diseases.  Prerequisites: BIO 221 or BIO 222 recommended. Offered in summer or January  term.
221 Human  Anatomy and Physiology I (4)
    An integrated lecture/laboratory  experience that examines the anatomy and physiology  of humans. Includes a survey of the nervous,  muscular, skeletal, integumentary, and respiratory systems.  Lecture three hours, laboratory three hours each week.
    $50 lab fee. Offered  every fall.
222 Human Anatomy and Physiology II (4)
    An integrated lecture/laboratory experience that examines the anatomy and physiology of humans. Includes  a survey of the  cardiovascular, endocrine, digestive, urinary and reproductive systems. Lecture  three hours, laboratory three hours each week. $50 lab fee. Offered every  spring.
 230/330 Urban Ecology (4)
    Investigation of urban  environments as ecosystems heavily influenced by human population and  consumption of natural resources. Examines impacts of urbanization, focusing on  issues related to water, air, energy and communicable diseases. Includes travel within and outside urban centers. Course fee  estimated at $5,000 required. Pass/fail grading. Prerequisite: BIO 130 or consent. Offered  intermittently.
 288 Genetics  (4)
    Principles of heredity as applied to both plants  and animals. Prerequisites: CHEM 120 and BIO 130 and 132. Lecture three hours, laboratory three hours each  week. Offered every spring.
301 Teaching  Experience (1)
    Qualified students  assist BIO instructors in the teaching  of biological sciences  courses and laboratories. May be repeated  for credit, but students may apply no more than 4 semester hours toward  graduation. Prerequisite: Consent.
316 General Ecology (4)
    A study of the interrelationships between organisms  and their environment. Topics will  range from the individual level to the  global scale, including both basic and applied  ecological topics. Prerequisites: BIO 130, successful completion of ENG 105  with a grade of C or higher, and sophomore status or higher. Identical  to EES 316. Offered every spring.
319 Field Studies in Coastal Environments (4)
    Couples ecological  principles with field observation  and data collection to understand the coastal environment. Major course topics include: organisms of  the marine coastal environments; sampling and analytical approaches for  studying plankton, invertebrates, nekton,  and plants; and conservation issues  in marine ecosystems. $200 lab fee. Offered every summer.
 320 Ornithology (4)
    A  study of birds  emphasizing diversity, behavioral ecology, and life history. Regular  field observations are required and may  occur under a variety of weather conditions. Prerequisite: BIO 130 and  sophomore status or higher. Lecture three hours, laboratory three hours each  week. Offered intermittently.
333 Conservation Biology  (4)
    This course  examines conservation at multiple levels  (genes to ecosystems) from a biological perspective. We examine what is currently known about  biodiversity loss, and the consequences of that  loss for ecosystems. Students will  build skills relevant to a conservation-related career, including review of  scientific evidence and management planning.
    Prerequisite: BIO 130, and sophomore status  or higher. Offered intermittently.
 340 Tropical  Marine Ecology (4)
    An introduction to major  tropical marine habitats and ecological  processes that control tropical marine ecosystems  with a focus on coral  reefs, mangroves and tropical seagrass  beds. Organisms characteristic of these habitats  and their symbiotic relationships will be examined.  Prerequisites: BIO 130/131 or BIO 200 or consent. Offered intermittently.
 355 Marine Ecology (4)
    A  study of organismal adaptation and community  organization in marine and estuarine habitats. A variety  of marine habitats are examined, with the laboratory  focusing primarily on local species and habitat types. Prerequisite: two  semesters of general biology or BIO 130 and sophomore status or higher or  consent. Offered every fall.
371 Histology (4)
    A detailed study of the cells, tissues, and organs that comprise the mammalian body. Intended for students seeking  careers in biology, medicine,  or veterinary sciences. Prerequisite: a grade of C or higher in BIO 132 and  sophomore status or higher.
    Lecture three hours, laboratory three hours each week. Offered  spring of off-numbered years.
372 Comparative Anatomy  (4)
    A  study of the evolution, morphology, and physiology of vertebrates. An intensive laboratory-directed examination of the major organ systems of vertebrates as  exemplified by the lamprey, dogfish, salamander, and cat. Intended for students  seeking careers in biology, medicine, or veterinary sciences. Prerequisites: BIO 200 with a grade of C or higher or consent.  Lecture three hours, laboratory three hours each week. Offered spring of  even-numbered years.
 373 Marine Invertebrate Zoology  (4)
    A survey of the marine invertebrate phyla, emphasizing the classification, evolution, ecology, morphology, and life histories of these organisms. Includes  laboratory examinations of representative groups and field sampling of local  invertebrate fauna.
    Prerequisite: BIO 200 or consent.  Lecture three hours,  laboratory three hours each week.  Offered fall of odd-numbered years.
375 Topics in Tropical  Biology (4)
    An intensive field experience in neotropical ecosystems (rainforests, coral reefs,  mangroves, caves, etc.). Descriptive studies  of local flora and fauna will be combined with an in-depth investigation  of a topic of interest. Field activities will include moderately strenuous  exercise under a variety of weather conditions. Destinations may include  Belize, Costa Rica, or other tropical sites. Course fee required. Identical to  EES 375. Prerequisite: BIO 130 and sophomore status or higher, and consent.
    Offered intermittently. Runs concurrently with BIO 435.
377 Entomology and Arachnology (4)
    A comprehensive survey of the  insects and arachnids of medical, economic, and ecological significance.  Includes laboratory examinations of living  and preserved specimens  and field collections of select taxa.  Prerequisite: BIO 200 or consent. Offered intermittently.
380 Comparative Animal Physiology (4)
    A  study of the basic mechanisms by which animals  function. Emphasizes how both invertebrate and vertebrate organisms change these basic mechanisms to  adapt to environmental conditions.  Prerequisites: a grade of C or higher in BIO 200 or consent. Lecture three  hours, laboratory three hours each week. Offered every spring.
 385 Animal Behavior (4)
    A study of the mechanisms and  evolution of animal behavior. Topics include genetics and development of  behavior, neural and physiological mechanisms of behavior, communication,  social behavior, habitat selection, reproductive behavior, and parental investment. Laboratory exercises provide  hands-on experiences for many of these concepts.  Prerequisites: BIO 130 and sophomore status and higher or consent. Lecture three hours, laboratory three hours each week. Offered every spring.
393 Scanning Electron  Microscopy (SEM) (2)
    Introduces the essentials and  broad research applications of SEM. Topics include the preparation of  biological and non- biological specimens, the use of secondary and backscattered electrons  for imaging, and the operation  of SEM in high vacuum and variable pressure modes.  Hands-on training and independent operation of the SEM are mandatory. Identical  to EES 393. Prerequisites: Sophomore status and consent. Offered  intermittently.
395 Fluorescence Microscopy (2)
    Examines the applications of epifluorescence and confocal laser microscopy to the imaging  of living cells and tissues.  Topics include sample preparation and a review of the principles of  fluorescence imaging. Practical training and independent operation of the  instrumentation are strongly emphasized. Prerequisites: Sophomore status and  consent. Offered intermittently.
 397 Advanced Light Microscopy (2)
    Reviews the fundamentals of light microscopy, with an emphasis  upon the compound  light microscope, stereomicroscope and polarizing light microscope. Topics include dark-field,  phase-contrast, differential interference contrast (DIC), simple fluorescence  and basic staining techniques. Use  of digital image capture and  manipulation software is required. Prerequisites: Sophomore status and  consent. Offered intermittently.
 399 Histotechnique (2)
    Introduces techniques used to prepare  tissues for light microscopy. Topics  include a review of laboratory safety procedures,  specimen fixation, tissue processing and embedding, sectioning with a  microtome, and staining. Hands-on training with a microtome is emphasized.  Prerequisites: Sophomore status and consent. Offered intermittently.
 410 Evolution (4)
    A study of the theory of  biological evolution through natural selection. Topics include micro evolution,  speciation, macroevolution, ecology and modern methods  of determining phylogenetic relationships. Prerequisites: BIO 130 and 132 and junior/senior status or consent.  Offered intermittently.
 412 Chemical Ecology (4)
    A study of how organisms use  chemicals to mediate interactions within and between species. Students will  gain experience interpreting primary research articles as varying topics are  discussed, such as plant-herbivore interactions, coral chemical defenses, and insect semiochemicals. Prerequisites: BIO 130, BIO 132, any 200-level CHEM course or consent. Lecture  three hours each week. Offered every fall.
 420 Cell and Molecular Biology (4)
    A study of the structure and  function of prokaryotes and eukaryotes at the cellular and molecular level.  Emphasizes the molecular nature of cellular structure, metabolism, and physiology. Prerequisite: BIO 288 (grade of C or higher). Offered every spring.
 424 Topics in Molecular  Techniques (2 or 4)
    A focused study on molecular  techniques necessary for research and biotechnology applications. Students’ conceptual understanding, troubleshooting and lab-skills will develop on given topics.  Topics vary with offerings and may include:  gene cloning, gene expression analysis or characterization of protein  expression and activity. May be  repeated as topics vary.
    Prerequisite: BIO 288 or consent.  Offered intermittently.
435 Tropical Ecology (4) I
    An intensive field experience in  neotropical ecosystems (coral reefs, rainforests, caves, mangrove swamps,  etc.), that links ecological phenomena with social and historical factors  to examine the conservation of biodiversity. Field  activities require moderately strenuous exercise and considerable hiking. Destinations may include Trinidad, Belize,  Costa Rica, or other tropical sites. Prerequisites: Students must have completed a lab science  course (L) and have junior or senior status. Course fee is additional. Student may take  this course one additional time for credit. Runs concurrently with BIO 375.
470 Internship in the Natural  Sciences (2 or 4)
    An intensive study of a specific  field of science  through an on-site  field experience with hands-on learning  opportunities that are relevant to the chosen site.  Students may enroll for 2 or 4 hours  in a given semester. A minimum of 80 hours devoted to the internship is expected for 2 semester hours, and a  minimum of 160 hours is expected for 4 semester hours, but some placements may  require more time. Students must coordinate their internship placement with the  supervising faculty member at least two months prior to placement. Pass/fail  grading. Prerequisites: junior/senior status and consent. Offered every  semester.
475 Natural  and Social History  of the Chesapeake Bay (4) I
    Provides a comprehensive view of  one of the largest and most diverse estuaries in the world. Students examine  the relationships between the natural history and the human history, including  social and political aspects, use of the bay by various societies and their impact on and preservation of the bay. Saturday field trips required.  Prerequisite: junior/senior  status. Offered intermittently.
 482 Microbiology (4)
    Teaches basic microbiological concepts  and the role of microorganisms in various applied areas. Topics include  microbial physiology, cell structure, microbial genetics, pathogenic  microorganisms and disease, and environmental and applied microbiology. Students  practice aseptic technique, isolation and identification of bacteria, staining, and determination of microbial numbers. Prerequisite: BIO  288 (grade of C or higher). Offered every fall.
489 Research in the Natural  Sciences (2 or 4)
    Offers students the opportunity to conduct original scientific research in an area of interest. Students  work closely with one or more members of the natural science  faculty to develop and conduct a research project, then present their findings  orally during the semester’s undergraduate research symposium and as a formal  research paper. Students are encouraged to present their findings at a  conference. Prerequisite:  junior/senior status and a major in  the natural sciences, prior approval by the project advisor, and  consent of the instructor. Students may enroll for 2 or 4 hours in a given  semester. May be repeated for a maximum total of 8 semester hours. Identical to  CHEM 489, CS 489, and EES 489. Offered  intermittently.